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Sara Shalin

TitleAssociate Professor
InstitutionUniversity of Arkansas for Medical Sciences
DepartmentPathology, College of Medicine
DivisionPathology Clinical
Address4S/12 Shorey
324 UAMS Campus Dr.
Mail Slot # 517
Little Rock AR 72205
Phone501-686-8007
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    Other Positions
    TitleAssociate Professor
    InstitutionUniversity of Arkansas for Medical Sciences
    DepartmentDermatology, College of Medicine


    Collapse Research Funds 
    Collapse research activities
    R01CA236209     (TACKETT, ALAN)Jun 15, 2019 - May 31, 2024
    NIH/Nat. Cancer Institute
    Identification of Druggable Targets to Complement Melanoma Therapy
    Role: Co-Investigator

    F31MH071014     (SHALIN, SARA C)Mar 1, 2004 - Jan 27, 2006
    NIH/NIMH
    Kinase Suppressor of ras: Function in the hippocampus
    Role: Principal Investigator

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    Collapse selected publications
    Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Faculty can login to make corrections and additions. Don't see publications published under other names? Login to add alternative names.
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    1. Singh AP, Jeffus SK, Shalin SC. Cervical dysplasia in a patient with inherited epidermodysplasia verruciformis-A mere coincidence? J Cutan Pathol. 2020 Dec 14. PMID: 33319409.
      View in: PubMed
    2. Jeffus SK, Quick CM, Chen C, Gardner JM, Kaley JR, Shalin SC. Prospective Consensus Reporting by Gynecologic Pathology and Dermatopathology Improves Diagnosis of Vulvar Biopsies. Arch Pathol Lab Med. 2020 11 19; 144(12):0. PMID: 33212498.
      View in: PubMed
    3. Dawson K, Bridge JA, Sumegi J, Royce T, Gardner JM, Shalin SC. Epithelioid Fibrous Histiocytoma With Dot-Like Perinuclear ALK Expression and PRKAR2A-ALK Fusion. Am J Dermatopathol. 2020 Nov; 42(11):861-864. PMID: 32379091.
      View in: PubMed
    4. Carlock S, Fulton EH, Rosenbaum E, Mukasa L, Gardner JM, Shalin SC, Kaley JR. Hansen Disease (Leprosy) and Armadillo Exposure in Arkansas: A Case Series. Am J Dermatopathol. 2020 Oct; 42(10):769-773. PMID: 32379089.
      View in: PubMed
    5. Campbell KK, Gardner JM, Hall K, Osborne A, Shalin SC. Consecutive Immunohistochemical Staining on a Single Slide is Effective in Confirming Melanocytic Derivation. Appl Immunohistochem Mol Morphol. 2020 10; 28(9):e76-e81. PMID: 30499815.
      View in: PubMed
    6. Taylor EM, Byrum SD, Edmondson JL, Wardell CP, Griffin BG, Shalin SC, Gokden M, Makhoul I, Tackett AJ, Rodriguez A. Proteogenomic analysis of melanoma brain metastases from distinct anatomical sites identifies pathways of metastatic progression. Acta Neuropathol Commun. 2020 09 05; 8(1):157. PMID: 32891176.
      View in: PubMed
    7. Shalin SC, Racher LM, Campbell KK. Lichenoid dermatoses involving the vulva: A clinical-pathologic correlation?. Semin Diagn Pathol. 2021 Jan; 38(1):3-18. PMID: 32951943.
      View in: PubMed
    8. Harrison DK, Waldrip ZJ, Burdine L, Shalin SC, Burdine MS. DNA-PKcs Inhibition Extends Allogeneic Skin Graft Survival. Transplantation. 2020 Sep 01. PMID: 32890138.
      View in: PubMed
    9. Shalin SC, Ferringer T, Cassarino DS. PAS and GMS utility in dermatopathology: Review of the current medical literature. J Cutan Pathol. 2020 Nov; 47(11):1096-1102. PMID: 32515092.
      View in: PubMed
    10. Fancher K, Gardner JM, Shalin SC. Elastophagocytosis and interstitial granulomatous infiltrate are more common in extragenital vs genital lichen sclerosus. J Cutan Pathol. 2020 Oct; 47(10):903-912. PMID: 32399972.
      View in: PubMed
    11. Donizy P, Wu CL, Mull J, Fujimoto M, Chlopik A, Peng Y, Shalin SC, Selim MA, Puig S, Fernandez-Figueras MT, Shea CR, Biernat W, Rys J, Marszalek A, Hoang MP. Up-Regulation of PARP1 Expression Significantly Correlated with Poor Survival in Mucosal Melanomas. Cells. 2020 05 05; 9(5). PMID: 32380691.
      View in: PubMed
    12. Cloutier JM, Shalin SC, Lindberg M, Gardner JM, Fernandez-Pol S, Zaba L, Novoa R, Brown RA. Cutaneous pleomorphic fibromas arising in patients with germline TP53 mutations. J Cutan Pathol. 2020 Aug; 47(8):734-741. PMID: 32187703.
      View in: PubMed
    13. Davis LE, Shalin SC, Tackett AJ. Utility of histone H3K27me3 and H4K20me as diagnostic indicators of melanoma. Melanoma Res. 2020 04; 30(2):159-165. PMID: 32142497.
      View in: PubMed
    14. Davis LE, Shalin SC, Tackett AJ. Current state of melanoma diagnosis and treatment. Cancer Biol Ther. 2019; 20(11):1366-1379. PMID: 31366280.
      View in: PubMed
    15. Shalin SC, Gardner JM, Kaley JR. Pearls and Updates in Dermatopathology for the General Surgical Pathologist: Experts Weigh In on What You Need to Know. Arch Pathol Lab Med. 2019 07; 143(7):786-788. PMID: 31225990.
      View in: PubMed
    16. Lora Gonzalez M, Webb M, Shalin S. Periumbilical Papule: Answer. Am J Dermatopathol. 2019 Jul; 41(7):531-532. PMID: 31232737.
      View in: PubMed
    17. Lora Gonzalez M, Webb M, Shalin S. Periumbilical Papule: Challenge. Am J Dermatopathol. 2019 Jul; 41(7):e71-e72. PMID: 29505474.
      View in: PubMed
    18. Wei T, Shalin S, Draper E, Miller E, Zhang H, Sun R, Lee M, Albert G, Richter GT. Abnormal elastin and collagen deposition is present in extracranial arteriovenous malformations: A comparison to intracranial disease. Histol Histopathol. 2019 Dec; 34(12):1355-1363. PMID: 31119718.
      View in: PubMed
    19. Abate MS, Battle LR, Emerson AN, Gardner JM, Shalin SC. Dermatologic Urgencies and Emergencies: What Every Pathologist Should Know. Arch Pathol Lab Med. 2019 08; 143(8):919-942. PMID: 30785787.
      View in: PubMed
    20. Shields BD, Koss B, Taylor EM, Storey AJ, West KL, Byrum SD, Mackintosh SG, Edmondson R, Mahmoud F, Shalin SC, Tackett AJ. Loss of E-Cadherin Inhibits CD103 Antitumor Activity and Reduces Checkpoint Blockade Responsiveness in Melanoma. Cancer Res. 2019 03 15; 79(6):1113-1123. PMID: 30674537.
      View in: PubMed
    21. Dunn ALJ, Gardner JM, Kaley JR, Bellamy W, Shalin SC. ALK Rearrangements Are Infrequent in Cellular Blue Nevus and Deep Penetrating Nevus. Am J Dermatopathol. 2018 Jul; 40(7):469-478. PMID: 29923908.
      View in: PubMed
    22. Abate MS, Shalin S. Progressive widespread telangiectasias. Cutis. 2018 May; 101(5):E1-E3. PMID: 29894536.
      View in: PubMed
    23. Dinehart M, Abate MS, Jennings T, Kaley JR, Shalin SC. Colliding, colonizing or combining? Four cases illustrating the unique challenges presented by melanoma arising in conjunction with basal cell carcinoma. J Cutan Pathol. 2018 Jun; 45(6):443-452. PMID: 29484694.
      View in: PubMed
    24. Purnell JC, Gardner JM, Brown JA, Shalin SC. Conventional Versus Giant Basal Cell Carcinoma, a Review of 57 Cases: Histologic Differences Contributing to Excessive Growth. Indian J Dermatol. 2018 Mar-Apr; 63(2):147-154. PMID: 29692457.
      View in: PubMed
    25. Haller J, David MP, Lee NE, Shalin SC, Gardner JM. Impact of Pathologist Involvement in Sarcoma and Rare Tumor Patient Support Groups on Facebook: A Survey of 542 Patients and Family Members. Arch Pathol Lab Med. 2018 09; 142(9):1113-1119. PMID: 29377717.
      View in: PubMed
    26. Chlopik A, Selim MA, Peng Y, Wu CL, Tell-Marti G, Paral KM, Shalin SC, Kraft S, Hsu CK, Shea CR, Puig S, Fernandez-Figueras MT, Biernat W, Rys J, Marszalek A, Hoang MP. Prognostic role of tumoral PDL1 expression and peritumoral FoxP3+ lymphocytes in vulvar melanomas. Hum Pathol. 2018 03; 73:176-183. PMID: 29307625.
      View in: PubMed
    27. Carlquist E, Lee NE, Shalin SC, Goodman M, Gardner JM. Dermatopathology and Social Media: Improving Participation by Increasing Clarity. Arch Pathol Lab Med. 2018 01; 142(1):11. PMID: 29257927.
      View in: PubMed
    28. Dunn A, Broadfoot BG, Hunt J, Kaley JR, Atiq O, Gutta NB, Wang X, Gardner JM, Shalin SC. Metastatic laryngeal large cell neuroendocrine carcinoma: A rare case of presentation and extreme tumor burden. J Cutan Pathol. 2018 Mar; 45(3):229-233. PMID: 29194700.
      View in: PubMed
    29. Dias-Santagata D, Selim MA, Su Y, Peng Y, Vollmer R, Chlopik A, Tell-Marti G, Paral KM, Shalin SC, Shea CR, Puig S, Fernandez-Figueras MT, Biernat W, Rys J, Marszalek A, Hoang MP. KIT mutations and CD117 overexpression are markers of better progression-free survival in vulvar melanomas. Br J Dermatol. 2017 Nov; 177(5):1376-1384. PMID: 28734009.
      View in: PubMed
    30. Durvasula VSPB, Shalin SC, Tulunay-Ugur OE, Suen JY, Richter GT. Effects of supramaximal balloon dilatation pressures on adult cricoid and tracheal cartilage: A cadaveric study. Laryngoscope. 2018 06; 128(6):1304-1309. PMID: 28988443.
      View in: PubMed
    31. Carlquist E, Lee NE, Shalin SC, Goodman M, Gardner JM. Dermatopathology and Social Media: A Survey of 131 Medical Professionals From 29 Countries. Arch Pathol Lab Med. 2018 Feb; 142(2):184-190. PMID: 28657771.
      View in: PubMed
    32. Liles JE, Shalin SC, White BA, Trigg LB, Kaley JR. Parvovirus B19 infection in an adult presenting with connective tissue disease-like symptoms: a report of the clinical and histological findings. Dermatol Online J. 2017 Jun 15; 23(6). PMID: 28633740.
      View in: PubMed
    33. Johnson G, Gardner JM, Shalin SC. Polarizable crystals in apocrine sweat gland tumors: A series of 3 cases. J Cutan Pathol. 2017 Aug; 44(8):698-702. PMID: 28497640.
      View in: PubMed
    34. Mahmoud F, Shields B, Makhoul I, Avaritt N, Wong HK, Hutchins LF, Shalin S, Tackett AJ. Immune surveillance in melanoma: From immune attack to melanoma escape and even counterattack. . 2017 Jul 03; 18(7):451-469. PMID: 28513269.
      View in: PubMed
    35. Shields BD, Mahmoud F, Taylor EM, Byrum SD, Sengupta D, Koss B, Baldini G, Ransom S, Cline K, Mackintosh SG, Edmondson RD, Shalin S, Tackett AJ. Indicators of responsiveness to immune checkpoint inhibitors. Sci Rep. 2017 04 11; 7(1):807. PMID: 28400597.
      View in: PubMed
    36. Kothari A, Shalin SC, Crescencio JC, Burgess MJ. Skin lesion in a patient with acute myeloid leukemia. Transpl Infect Dis. 2017 Feb; 19(1). PMID: 27862750.
      View in: PubMed
    37. Shalin SC. A review of kinase fusions in melanocytic tumors. Lab Invest. 2017 02; 97(2):158-165. PMID: 27892929.
      View in: PubMed
    38. Romano RC, Gardner JM, Shalin SC, Ram R, Govindarajan R, Montgomery CO, Gilley JH, Nicholas RW. High Relative Expression of Pannexin 3 (PANX3) in an Axillary Sweat Gland Carcinoma With Osteosarcomatous Transformation. Am J Dermatopathol. 2016 Nov; 38(11):846-851. PMID: 27391460.
      View in: PubMed
    39. Kaliki V, Shalin SC, Edgar-Zarate C. A Rare Rash in a Pediatric Patient Due to a Very Common Medication. Clin Pediatr (Phila). 2017 Jan; 56(1):97-99. PMID: 27029819.
      View in: PubMed
    40. Kothari A, Shalin SC, Rico Crescencio JC, Burgess MJ. Skin Lesion with Splenic Microabscesses in a Patient with Acute Myeloid Leukemia. Am J Med. 2016 Dec; 129(12):e325-e327. PMID: 27637597.
      View in: PubMed
    41. Mohan M, Shalin SC, Kothari A, Rico JC, Caradine K, Burgess M. Lasiodiplodia species fungal osteomyelitis in a multiple myeloma patient. Transpl Infect Dis. 2016 Oct; 18(5):761-764. PMID: 27391908.
      View in: PubMed
    42. Kaur V, Swami A, Shebli A, Shalin S, Veeraputhiran M, Emanuel P, Jethava Y. A rare case of blastic plasmacytoid dendritic cell neoplasm with deletion 7q.31, in the setting of heavy pre-treatment with alkylating chemotherapy. J Oncol Pharm Pract. 2017 Oct; 23(7):552-556. PMID: 27559020.
      View in: PubMed
    43. Jones C, Shalin SC, Gardner JM. Incidence of mature adipocytic component within cutaneous smooth muscle neoplasms. J Cutan Pathol. 2016 Oct; 43(10):866-71. PMID: 27404774.
      View in: PubMed
    44. Erstine EM, Elwood HR, Westbrook KC, McCalmont TH, Shalin SC, Gardner JM. Desmoplastic melanoma presenting as primary alopecia neoplastica: a report of two cases. J Cutan Pathol. 2016 Oct; 43(10):872-9. PMID: 27272456.
      View in: PubMed
    45. Shields BD, Tackett AJ, Shalin SC. Proteomics and melanoma: a current perspective. Glob Dermatol. 2016 Aug; 3(4):366-370. PMID: 30214824.
      View in: PubMed
    46. Clark LN, Elwood HR, Uhlenhake EE, Smoller BR, Shalin SC, Gardner JM. Nuclear factor XIIIa staining (clone AC-1A1 mouse monoclonal) is a highly sensitive marker of sebaceous differentiation in normal and neoplastic sebocytes. J Cutan Pathol. 2016 Aug; 43(8):657-62. PMID: 27153463.
      View in: PubMed
    47. Uhlenhake EE, Clark LN, Smoller BR, Shalin SC, Gardner JM. Nuclear factor XIIIa staining (clone AC-1A1 mouse monoclonal) is a sensitive and specific marker to discriminate sebaceous proliferations from other cutaneous clear cell neoplasms. J Cutan Pathol. 2016 Aug; 43(8):649-56. PMID: 27153339.
      View in: PubMed
    48. Fulton E, Alapat D, Mabry A, Gao L, Shalin SC. CLL/SLL with initial presentation as a longstanding cutaneous plaque: a rare case. J Cutan Pathol. 2016 Aug; 43(8):717-21. PMID: 27147484.
      View in: PubMed
    49. Schnebelen AM, Gardner JM, Shalin SC. Margin Status in Shave Biopsies of Nonmelanoma Skin Cancers: Is It Worth Reporting? Arch Pathol Lab Med. 2016 Jul; 140(7):678-81. PMID: 27116089.
      View in: PubMed
    50. Mahmoud F, Shields B, Makhoul I, Hutchins LF, Shalin SC, Tackett AJ. Role of EZH2 histone methyltrasferase in melanoma progression and metastasis. . 2016 06 02; 17(6):579-91. PMID: 27105109.
      View in: PubMed
    51. Uhlenhake EE, Smoller BR, Elwood HR, Shalin SC, Gardner JM. Characterization of dermatopathology fellowship applicants: a 5-year single institution experience. J Cutan Pathol. 2016 May; 43(5):411-7. PMID: 26863905.
      View in: PubMed
    52. Purnell JC, Williams BA, Shalin SC, Wong HK. Mucocutaneous findings associated with interleukin (IL)-17 inhibition. JAAD Case Rep. 2016 Jan; 2(1):92-4. PMID: 27051840.
      View in: PubMed
    53. Shields B, Shalin SC, Tackett AJ. Microscopes and Mass Spectrometers. J Proteomics Bioinform. 2016; 9(Suppl 10). PMID: 29910559.
      View in: PubMed
    54. Sengupta D, Byrum SD, Avaritt NL, Davis L, Shields B, Mahmoud F, Reynolds M, Orr LM, Mackintosh SG, Shalin SC, Tackett AJ. Quantitative Histone Mass Spectrometry Identifies Elevated Histone H3 Lysine 27 (Lys27) Trimethylation in Melanoma. Mol Cell Proteomics. 2016 Mar; 15(3):765-75. PMID: 26621846.
      View in: PubMed
    55. Kaskas NM, Kern M, Johnson A, Hughes MP, Hardee M, Day J, Gokden M, Shalin SC, Gao L. A Case of Cutaneous Rosai-Dorfman Disease with underlying calvarial involvement and absence of BRAFV600E mutation. JAAD Case Rep. 2015 Nov 01; 1(6):408-411. PMID: 26858984.
      View in: PubMed
    56. Elwood H, Berry RS, Gardner JM, Shalin SC. Superficial fibrin thrombi … and other findings: a review of the histopathology of human scabietic infections. J Cutan Pathol. 2015 May; 42(5):346-52. PMID: 25754497.
      View in: PubMed
    57. Harms PW, Fullen DR, Patel RM, Chang D, Shalin SC, Ma L, Wood B, Beer TW, Siddiqui J, Carskadon S, Wang M, Palanisamy N, Fisher GJ, Andea A. Cutaneous basal cell carcinosarcomas: evidence of clonality and recurrent chromosomal losses. Hum Pathol. 2015 May; 46(5):690-7. PMID: 25704628.
      View in: PubMed
    58. Cole AJ, Chen C, Lorsbach RB, Honnebier BM, Gardner JM, Shalin SC. Extranodal Rosai-Dorfman disease in the scrotum of a 13-month male: a unique anatomic presentation. Am J Dermatopathol. 2015 Jan; 37(1):88-90. PMID: 24999545.
      View in: PubMed
    59. Schnebelen AM, Page J, Gardner JM, Shalin SC. Benign lymphangioendothelioma presenting as a giant flank mass. J Cutan Pathol. 2015 Mar; 42(3):217-21. PMID: 25410589.
      View in: PubMed
    60. Uhlenhake EE, Smoller BR, Gardner JM, Shalin SC. Epithelioid melanocytic nevus with tubule and pseudoacini formation. J Cutan Pathol. 2015 Mar; 42(3):213-6. PMID: 25407962.
      View in: PubMed
    61. Battle LR, Shalin SC, Gao L. Granulomatous pigmented purpuric dermatosis. Clin Exp Dermatol. 2015 Jun; 40(4):387-90. PMID: 25522174.
      View in: PubMed
    62. Johnson M, Walker D, Galloway W, Gardner JM, Shalin SC. Interface dermatitis along Blaschko's lines. J Cutan Pathol. 2014 Dec; 41(12):950-4. PMID: 25353350.
      View in: PubMed
    63. Shalin SC, Cifarelli CP, Suen JY, Gao L. Loss of cytokeratin 20 and acquisition of thyroid transcription factor-1 expression in a merkel cell carcinoma metastasis to the brain. Am J Dermatopathol. 2014 Nov; 36(11):904-6. PMID: 24901475.
      View in: PubMed
    64. Stuart LN, Hiatt KM, Zaki Z, Gardner JM, Shalin SC. Plaque-like CD34-positive dermal fibroma/medallion-like dermal dendrocyte hamartoma: an unusual spindle cell neoplasm. J Cutan Pathol. 2014 Aug; 41(8):625-9. PMID: 25065391.
      View in: PubMed
    65. Lataifeh AR, Deas S, Shalin SC, Khasawneh KR. A 62-year-old women with persistent severe asthma, skin rash, and eosinophilia. Chest. 2014 Aug; 146(2):e52-e55. PMID: 25091763.
      View in: PubMed
    66. Elwood H, Gardner J, Galloway W, Shalin SC. A case of a giant sarcomatoid basal cell carcinoma with heterologous osteosarcomatous elements. J Cutan Pathol. 2014 Jun; 41(6):550-1. PMID: 24684278.
      View in: PubMed
    67. Shalin SC, Rinaldi C, Horn TD. Clear cell acanthoma with changes of eccrine syringofibroadenoma: reactive change or clue to etiology? J Cutan Pathol. 2013 Dec; 40(12):1021-6. PMID: 24117902.
      View in: PubMed
    68. McDermott A, Jesus AA, Liu Y, Kim P, Jacks J, Montealegre Sanchez GA, Chen Y, Kannan A, Schnebelen A, Emanuel PD, Shalin S, Hiatt K, Goldbach-Mansky R, Gao L. A case of proteasome-associated auto-inflammatory syndrome with compound heterozygous mutations. J Am Acad Dermatol. 2013 Jul; 69(1):e29-32. PMID: 23768303.
      View in: PubMed
    69. Shalin SC, Haws AL, Carter DG, Zarrin-Khameh N. Clear cell adenocarcinoma arising from endometriosis in abdominal wall cesarean section scar: a case report and review of the literature. J Cutan Pathol. 2012 Nov; 39(11):1035-41. PMID: 22882475.
      View in: PubMed
    70. Shalin SC, Brantley J, Diwan AH. Follicular mucinosis and mycosis-fungoides-like drug eruption due to leuprolide acetate: a case report and review. J Cutan Pathol. 2012 Nov; 39(11):1022-5. PMID: 22882386.
      View in: PubMed
    71. Shalin SC, Sakharpe A, Lyle S, Lev D, Calonje E, Lazar AJ. p53 staining correlates with tumor type and location in sebaceous neoplasms. Am J Dermatopathol. 2012 Apr; 34(2):129-35; quiz 136-8. PMID: 22441365.
      View in: PubMed
    72. Shalin SC, Lyle S, Calonje E, Lazar AJ. Sebaceous neoplasia and the Muir-Torre syndrome: important connections with clinical implications. Histopathology. 2010 Jan; 56(1):133-47. PMID: 20055911.
      View in: PubMed
    73. Shalin SC, Hernandez CM, Dougherty MK, Morrison DK, Sweatt JD. Kinase suppressor of Ras1 compartmentalizes hippocampal signal transduction and subserves synaptic plasticity and memory formation. Neuron. 2006 Jun 01; 50(5):765-79. PMID: 16731514.
      View in: PubMed
    74. Shalin SC, Egli R, Birnbaum SG, Roth TL, Levenson JM, Sweatt JD. Signal transduction mechanisms in memory disorders. Prog Brain Res. 2006; 157:25-41. PMID: 17167902.
      View in: PubMed
    75. Shalin. A role for kinase supporessor of ras (KSR1) in the hippocampus. 2005.
    76. Shalin SC, Zirrgiebel U, Honsa KJ, Julien JP, Miller FD, Kaplan DR, Sweatt JD. Neuronal MEK is important for normal fear conditioning in mice. J Neurosci Res. 2004 Mar 15; 75(6):760-70. PMID: 14994337.
      View in: PubMed
    77. Shalin. Kinase supporessor of ras facilitates activation of the ERK/MAP kinase cascade within the hippocampus and is important for learning. 2004.
    78. Shalin. Neuronal specific dominant negative MEK produces specific contextual fear conditioning alterations depending on the dosage of the transgene. 2003.
    79. Shalin. Kinase suppressor of ras facilitates activation of the ERK/MAP kinase cascade within the hippocampus and is important for learning. 2003.
    80. Shalin S. Kinase supporessor of ras facilitates activation of the ERK/MAP kinase cascade within the hippocampus and may be important for learning. 2003.
    81. Niswender CM, Shalin, Herrick-Davis K, Emeson RB, Sanders-Bush E. RNA editing of the human serotonin 5-hydroxytryptamine 2C receptor slicences constitutive activity. Journal of Biological Chemistry. 1999; 274(14):9472-9478.
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